A Galápagos tortoise named Gramma died at the San Diego Zoo on November 20, 2025, after an estimated lifespan of about 141 years. Records showed that the tortoise originally came from the Galápagos Islands and later arrived from the Bronx Zoo, possibly in 1928 or 1931, for an early conservation program. Gramma lived through major global events, including two world wars and the terms of 20 United States presidents. Zoo specialists said the animal developed severe age-related bone problems that worsened in recent months, and the tortoise was euthanized to prevent additional suffering. Many visitors shared memories of seeing Gramma, and longtime guests described the animal as a symbol of endurance and natural heritage. Experts noted that Galápagos tortoises usually live more than 100 years in the wild and can reach nearly double that age with human care.

Veterinarians reported that Gramma received specialized support during the final stage of life, including monitoring for skeletal changes often seen in elderly tortoises. The zoo also described the wider conservation situation. Galápagos tortoises include 15 subspecies, with three already extinct and the remaining groups listed as vulnerable or critically endangered by the International Union for Conservation of Nature. Breeding programs have released more than 10,000 young tortoises into natural habitats since 1965 to rebuild populations. Recent hatchings at two United States zoos showed that even very old adults can still reproduce. Scientists said ongoing protection, targeted research, and careful habitat management remain essential for long-term survival.