A long-lived Galápagos tortoise named Gramma died at the San Diego Zoo on November 20, 2025, after an estimated lifespan of about 141 years. Zoo records showed that the tortoise originally came from the Galápagos Islands and later arrived from the Bronx Zoo, possibly in 1928 or 1931, as part of an early conservation group. Gramma lived through major historical periods, including two global wars and the terms of 20 United States presidents. Zoo specialists stated that the animal developed serious age-related bone conditions that recently worsened, leading to euthanasia. Many visitors shared memories of early encounters with the tortoise, and several longtime guests described Gramma as a symbol of endurance and natural heritage. Experts explained that Galápagos tortoises usually live for more than 100 years in the wild and can reach close to double that age in human care, making this species an emblematic example of extreme longevity.

Veterinarians reported that Gramma received specialized support during the final stage of life, including monitoring for skeletal changes that often affect elderly tortoises. The zoo also highlighted the wider conservation context, noting that Galápagos tortoises consist of 15 subspecies, with three considered extinct and the rest classified as vulnerable or critically endangered by the International Union for Conservation of Nature. Breeding programs have released more than 10,000 juveniles into natural habitats since 1965, demonstrating a significant and deliberate effort to restore declining populations. Recent hatchings at the Philadelphia Zoo and Zoo Miami showed continued reproductive success among extremely old adults, a development viewed as encouraging for future recovery. Scientists emphasized that ongoing protection, genetic research, and habitat management remain crucial for long-term species survival, despite persistent ecological threats and habitat fragmentation.